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Seekers run along the ground, spreading a narrow linear line of ink behind them - swim through this ink trail to quickly get around a stage. In their default mode, Seekers continue moving in a straight line until they hit an object, at which point they explode. But Seekers can also lock on to opponents. Hold down the sub weapon button to lock on to visible opponents - a reticle appears over your target. Release the sub weapon button to send the Seeker toward your target. While Seekers will attempt to chase down your target, they're incapable of making sharp turns or traversing steep inclines. But they can zoom right up most ramps, perfect for surprising opponents on the other side.
Splatoon Prima Guide